Potential Gaps in catabolism of small carbon sources in Weissella oryzae SG25
Found 152 low-confidence and 26 medium-confidence steps on the best paths for 62 pathways.
Pathway | Step | Best candidate | 2nd candidate |
2-oxoglutarate | kgtP: 2-oxoglutarate:H+ symporter KgtP | | |
4-hydroxybenzoate | mhpD: 2-hydroxypentadienoate hydratase | | |
4-hydroxybenzoate | mhpE: 4-hydroxy-2-oxovalerate aldolase | | |
4-hydroxybenzoate | pcaK: 4-hydroxybenzoate transporter pcaK | | |
4-hydroxybenzoate | pobA: 4-hydroxybenzoate 3-monooxygenase | | |
4-hydroxybenzoate | praA: protocatechuate 2,3-dioxygenase | | |
4-hydroxybenzoate | xylF: 2-hydroxymuconate semialdehyde hydrolase | | |
acetate | ybhL: acetate uptake transporter YbhL | TT13_RS05825 | |
arginine | rocA: 1-pyrroline-5-carboxylate dehydrogenase | | |
arginine | rocD: ornithine aminotransferase | TT13_RS00935 | |
arginine | rocE: L-arginine permease | TT13_RS08760 | TT13_RS01405 |
asparagine | ansP: L-asparagine permease AnsP | TT13_RS01405 | TT13_RS00995 |
aspartate | glt: aspartate:proton symporter Glt | | |
cellobiose | bgl: cellobiase | | |
cellobiose | manY: glucose PTS, enzyme EIIC | TT13_RS09610 | |
citrate | SLC13A5: citrate:Na+ symporter | | |
citrulline | AO353_03040: ABC transporter for L-Citrulline, ATPase component | TT13_RS07590 | TT13_RS01035 |
citrulline | AO353_03045: ABC transporter for L-Citrulline, permease component 2 | TT13_RS01030 | |
citrulline | AO353_03050: ABC transporter for L-Citrulline, permease component 1 | TT13_RS07595 | TT13_RS01030 |
citrulline | AO353_03055: ABC transporter for L-Citrulline, periplasmic substrate-binding component | | |
citrulline | rocA: 1-pyrroline-5-carboxylate dehydrogenase | | |
citrulline | rocD: ornithine aminotransferase | TT13_RS00935 | |
D-alanine | dadA: D-alanine dehydrogenase | | |
D-serine | dsdA: D-serine ammonia-lyase | | |
deoxyinosine | deoB: phosphopentomutase | TT13_RS03255 | |
deoxyinosine | deoC: deoxyribose-5-phosphate aldolase | | |
deoxyinosine | deoD: deoxyinosine phosphorylase | | |
deoxyribonate | aacS: acetoacetyl-CoA synthetase | | |
deoxyribonate | deoxyribonate-dehyd: 2-deoxy-D-ribonate 3-dehydrogenase | TT13_RS08720 | |
deoxyribonate | deoxyribonate-transport: 2-deoxy-D-ribonate transporter | | |
deoxyribonate | garK: glycerate 2-kinase | | |
deoxyribonate | ketodeoxyribonate-cleavage: 2-deoxy-3-keto-D-ribonate cleavage enzyme | | |
deoxyribose | deoC: deoxyribose-5-phosphate aldolase | | |
deoxyribose | deoP: deoxyribose transporter | | |
fructose | levF: fructose PTS system (fructose 6-phosphate forming), EII-C component | TT13_RS09610 | |
fructose | levG: fructose PTS system (fructose 6-phosphate forming), EII-D component | TT13_RS09605 | |
fucose | aldA: lactaldehyde dehydrogenase | | |
fucose | fucA: L-fuculose-phosphate aldolase FucA | | |
fucose | fucI: L-fucose isomerase FucI | | |
fucose | fucK: L-fuculose kinase FucK | | |
fucose | fucP: L-fucose:H+ symporter FucP | | |
fucose | fucU: L-fucose mutarotase FucU | | |
fumarate | dctA: fumarate:H+ symporter DctA | | |
galactose | galP: galactose:H+ symporter GalP | TT13_RS07120 | TT13_RS06920 |
galacturonate | eda: 2-keto-3-deoxygluconate 6-phosphate aldolase | | |
galacturonate | exuT: D-galacturonate transporter ExuT | | |
galacturonate | uxaA: D-altronate dehydratase | | |
galacturonate | uxaB: tagaturonate reductase | | |
galacturonate | uxaC: D-galacturonate isomerase | | |
glucosamine | manX: glucosamine PTS system, EII-AB component ManX | TT13_RS09615 | |
glucosamine | manY: glucosamine PTS system, EII-C component ManY | TT13_RS09610 | |
glucosamine | manZ: glucosamine PTS system, EII-D component ManZ | TT13_RS09605 | |
glucose | manY: glucose PTS, enzyme EIIC | TT13_RS09610 | |
glucose-6-P | uhpT: glucose-6-phosphate:phosphate antiporter | | |
glucuronate | eda: 2-keto-3-deoxygluconate 6-phosphate aldolase | | |
glucuronate | exuT: D-glucuronate:H+ symporter ExuT | | |
glucuronate | uxaC: D-glucuronate isomerase | | |
glucuronate | uxuA: D-mannonate dehydratase | | |
glucuronate | uxuB: D-mannonate dehydrogenase | | |
glutamate | gdhA: glutamate dehydrogenase, NAD-dependent | | |
glutamate | glnP: L-glutamate ABC transporter, fused permease and substrate-binding components GlnP | | |
glycerol | glpD: glycerol 3-phosphate dehydrogenase (monomeric) | | |
histidine | hutG: N-formiminoglutamate formiminohydrolase | | |
histidine | hutH: histidine ammonia-lyase | | |
histidine | hutI: imidazole-5-propionate hydrolase | | |
histidine | hutU: urocanase | | |
isoleucine | acdH: (2S)-2-methylbutanoyl-CoA dehydrogenase | | |
isoleucine | ech: 2-methyl-3-hydroxybutyryl-CoA hydro-lyase | | |
isoleucine | epi: methylmalonyl-CoA epimerase | | |
isoleucine | ivdG: 3-hydroxy-2-methylbutyryl-CoA dehydrogenase | TT13_RS08720 | |
isoleucine | mcmA: methylmalonyl-CoA mutase, fused catalytic and adenosylcobamide-binding components | | |
isoleucine | ofo: branched-chain alpha-ketoacid:ferredoxin oxidoreductase, fused | | |
isoleucine | pccA: propionyl-CoA carboxylase, alpha subunit | TT13_RS08740 | |
isoleucine | pccB: propionyl-CoA carboxylase, beta subunit | | |
L-lactate | lctO: L-lactate oxidase or 2-monooxygenase | | |
lactose | lacS: lactose permease LacS | TT13_RS08900 | |
leucine | aacS: acetoacetyl-CoA synthetase | | |
leucine | ilvE: L-leucine transaminase | TT13_RS06065 | |
leucine | liuA: isovaleryl-CoA dehydrogenase | | |
leucine | liuB: 3-methylcrotonyl-CoA carboxylase, alpha (biotin-containing) subunit | TT13_RS08740 | |
leucine | liuC: 3-methylglutaconyl-CoA hydratase | | |
leucine | liuD: 3-methylcrotonyl-CoA carboxylase, beta subunit | | |
leucine | liuE: hydroxymethylglutaryl-CoA lyase | | |
leucine | ofo: branched-chain alpha-ketoacid:ferredoxin oxidoreductase, fused | | |
lysine | amaB: L-2-aminoadipate semialdehyde dehydrogenase (AmaB/Pcd) | | |
lysine | hglS: D-2-hydroxyglutarate synthase | | |
lysine | lat: L-lysine 6-aminotransferase | TT13_RS00935 | |
lysine | lysN: 2-aminoadipate transaminase | TT13_RS06065 | |
lysine | lysP: L-lysine:H+ symporter LysP | TT13_RS08760 | TT13_RS01405 |
lysine | ydiJ: (R)-2-hydroxyglutarate dehydrogenase | | |
maltose | malI: maltose transporter | TT13_RS01885 | |
mannitol | mt2d: mannitol 2-dehydrogenase | TT13_RS08720 | TT13_RS05830 |
mannitol | PLT5: polyol transporter PLT5 | TT13_RS07120 | TT13_RS06920 |
myoinositol | iolB: 5-deoxy-D-glucuronate isomerase | | |
myoinositol | iolC: 5-dehydro-2-deoxy-D-gluconate kinase | | |
myoinositol | iolD: 3D-(3,5/4)-trihydroxycyclohexane-1,2-dione hydrolase | | |
myoinositol | iolE: scyllo-inosose 2-dehydratase | | |
myoinositol | iolG: myo-inositol 2-dehydrogenase | | |
myoinositol | iolJ: 5-dehydro-2-deoxyphosphogluconate aldolase | TT13_RS08845 | |
myoinositol | iolT: myo-inositol:H+ symporter | TT13_RS06920 | TT13_RS07120 |
myoinositol | mmsA: malonate-semialdehyde dehydrogenase | | |
NAG | nagEcba: N-acetylglucosamine phosphotransferase system, EII-CBA components | | |
phenylacetate | H281DRAFT_04042: phenylacetate:H+ symporter | TT13_RS00995 | TT13_RS01405 |
phenylacetate | paaA: phenylacetyl-CoA 1,2-epoxidase, subunit A | | |
phenylacetate | paaB: phenylacetyl-CoA 1,2-epoxidase, subunit B | | |
phenylacetate | paaC: phenylacetyl-CoA 1,2-epoxidase, subunit C | | |
phenylacetate | paaE: phenylacetyl-CoA 1,2-epoxidase, subunit E | | |
phenylacetate | paaF: 2,3-dehydroadipyl-CoA hydratase | | |
phenylacetate | paaG: 1,2-epoxyphenylacetyl-CoA isomerase / 2-(oxepinyl)acetyl-CoA isomerase / didehydroadipyl-CoA isomerase | | |
phenylacetate | paaH: 3-hydroxyadipyl-CoA dehydrogenase | TT13_RS08720 | |
phenylacetate | paaJ1: 3-oxo-5,6-dehydrosuberyl-CoA thiolase | TT13_RS02270 | |
phenylacetate | paaJ2: 3-oxoadipyl-CoA thiolase | TT13_RS02270 | |
phenylacetate | paaK: phenylacetate-CoA ligase | | |
phenylacetate | paaZ1: oxepin-CoA hydrolase | | |
phenylacetate | paaZ2: 3-oxo-5,6-didehydrosuberyl-CoA semialdehyde dehydrogenase | | |
phenylalanine | aacS: acetoacetyl-CoA synthetase | | |
phenylalanine | fahA: fumarylacetoacetate hydrolase | | |
phenylalanine | hmgA: homogentisate dioxygenase | | |
phenylalanine | HPD: 4-hydroxyphenylpyruvate dioxygenase | | |
phenylalanine | maiA: maleylacetoacetate isomerase | | |
phenylalanine | PAH: phenylalanine 4-monooxygenase | | |
phenylalanine | PCBD: pterin-4-alpha-carbinoalamine dehydratase | | |
phenylalanine | QDPR: 6,7-dihydropteridine reductase | | |
proline | put1: proline dehydrogenase | | |
proline | putA: L-glutamate 5-semialdeyde dehydrogenase | | |
propionate | epi: methylmalonyl-CoA epimerase | | |
propionate | mcmA: methylmalonyl-CoA mutase, fused catalytic and adenosylcobamide-binding components | | |
propionate | pccA: propionyl-CoA carboxylase, alpha subunit | TT13_RS08740 | |
propionate | pccB: propionyl-CoA carboxylase, beta subunit | | |
propionate | prpE: propionyl-CoA synthetase | | |
propionate | putP: propionate transporter; proline:Na+ symporter | | |
putrescine | gabD: succinate semialdehyde dehydrogenase | TT13_RS05060 | |
putrescine | gabT: gamma-aminobutyrate transaminase | TT13_RS00935 | |
putrescine | patA: putrescine aminotransferase (PatA/SpuC) | TT13_RS00935 | |
putrescine | patD: gamma-aminobutyraldehyde dehydrogenase | | |
pyruvate | SLC5A8: sodium-coupled pyruvate transporter | | |
rhamnose | aldA: lactaldehyde dehydrogenase | | |
rhamnose | rhaA: L-rhamnose isomerase | | |
rhamnose | rhaB: L-rhamnulokinase | | |
rhamnose | rhaD: rhamnulose 1-phosphate aldolase | | |
rhamnose | rhaM: L-rhamnose mutarotase | | |
rhamnose | rhaT: L-rhamnose:H+ symporter RhaT | | |
serine | sdaB: L-serine ammonia-lyase | | |
serine | serP: L-serine permease SerP | TT13_RS00995 | TT13_RS01405 |
sorbitol | sdh: sorbitol dehydrogenase | TT13_RS08680 | TT13_RS08720 |
sorbitol | SOT: sorbitol:H+ co-transporter SOT1 or SOT2 | TT13_RS07120 | TT13_RS06920 |
succinate | sdc: succinate:Na+ symporter Sdc | | |
sucrose | ams: sucrose hydrolase (invertase) | | |
sucrose | manY: glucose PTS, enzyme EIIC | TT13_RS09610 | |
threonine | gcvH: glycine cleavage system, H component (lipoyl protein) | | |
threonine | gcvP: glycine cleavage system, P component (glycine decarboxylase) | | |
threonine | gcvT: glycine cleavage system, T component (tetrahydrofolate aminomethyltransferase) | | |
threonine | ltaE: L-threonine aldolase | TT13_RS00625 | |
threonine | serP1: L-threonine uptake transporter SerP1 | TT13_RS00995 | TT13_RS01405 |
thymidine | deoA: thymidine phosphorylase DeoA | | |
thymidine | deoB: phosphopentomutase | TT13_RS03255 | |
thymidine | deoC: deoxyribose-5-phosphate aldolase | | |
thymidine | nupG: thymidine permease NupG/XapB | | |
trehalose | manY: glucose PTS, enzyme EIIC | TT13_RS09610 | |
trehalose | treF: trehalase | | |
tryptophan | tnaA: tryptophanase | | |
tyrosine | aacS: acetoacetyl-CoA synthetase | | |
tyrosine | fahA: fumarylacetoacetate hydrolase | | |
tyrosine | hmgA: homogentisate dioxygenase | | |
tyrosine | HPD: 4-hydroxyphenylpyruvate dioxygenase | | |
tyrosine | maiA: maleylacetoacetate isomerase | | |
valine | acdH: isobutyryl-CoA dehydrogenase | | |
valine | bch: 3-hydroxyisobutyryl-CoA hydrolase | | |
valine | ech: (S)-3-hydroxybutanoyl-CoA hydro-lyase | | |
valine | epi: methylmalonyl-CoA epimerase | | |
valine | mcmA: methylmalonyl-CoA mutase, fused catalytic and adenosylcobamide-binding components | | |
valine | mmsA: methylmalonate-semialdehyde dehydrogenase | | |
valine | mmsB: 3-hydroxyisobutyrate dehydrogenase | TT13_RS05300 | |
valine | ofo: branched-chain alpha-ketoacid:ferredoxin oxidoreductase, fused | | |
valine | pccA: propionyl-CoA carboxylase, alpha subunit | TT13_RS08740 | |
valine | pccB: propionyl-CoA carboxylase, beta subunit | | |
xylitol | PLT5: xylitol:H+ symporter PLT5 | TT13_RS07120 | TT13_RS06920 |
xylitol | xdhA: xylitol dehydrogenase | TT13_RS08545 | TT13_RS05830 |
Confidence: high confidence medium confidence low confidence
This GapMind analysis is from Sep 24 2021. The underlying query database was built on Sep 17 2021.
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About GapMind
Each pathway is defined by a set of rules based on individual steps or genes. Candidates for each step are identified by using
ublast (a fast alternative to protein BLAST)
against a database of manually-curated proteins (most of which are experimentally characterized) or by using
HMMer with enzyme models (usually from
TIGRFam). Ublast hits may be split across two different proteins.
A candidate for a step is "high confidence" if either:
- ublast finds a hit to a characterized protein at above 40% identity and 80% coverage, and bits >= other bits+10.
- (Hits to curated proteins without experimental data as to their function are never considered high confidence.)
- HMMer finds a hit with 80% coverage of the model, and either other identity < 40 or other coverage < 0.75.
where "other" refers to the best ublast hit to a sequence that is not annotated as performing this step (and is not "ignored").
Otherwise, a candidate is "medium confidence" if either:
- ublast finds a hit at above 40% identity and 70% coverage (ignoring otherBits).
- ublast finds a hit at above 30% identity and 80% coverage, and bits >= other bits.
- HMMer finds a hit (regardless of coverage or other bits).
Other blast hits with at least 50% coverage are "low confidence."
Steps with no high- or medium-confidence candidates may be considered "gaps."
For the typical bacterium that can make all 20 amino acids, there are 1-2 gaps in amino acid biosynthesis pathways.
For diverse bacteria and archaea that can utilize a carbon source, there is a complete
high-confidence catabolic pathway (including a transporter) just 38% of the time, and
there is a complete medium-confidence pathway 63% of the time.
Gaps may be due to:
- our ignorance of proteins' functions,
- omissions in the gene models,
- frame-shift errors in the genome sequence, or
- the organism lacks the pathway.
GapMind relies on the predicted proteins in the genome and does not search the six-frame translation. In most cases, you can search the six-frame translation by clicking on links to Curated BLAST for each step definition (in the per-step page).
For more information, see:
If you notice any errors or omissions in the step descriptions, or any questionable results, please let us know
by Morgan Price, Arkin group, Lawrence Berkeley National Laboratory