Potential Gaps in catabolism of small carbon sources in Erythrobacter gangjinensis K7-2
Found 114 low-confidence and 55 medium-confidence steps on the best paths for 62 pathways.
Pathway | Step | Best candidate | 2nd candidate |
2-oxoglutarate | kgtP: 2-oxoglutarate:H+ symporter KgtP | AAW01_RS02550 | |
4-hydroxybenzoate | adh: acetaldehyde dehydrogenase (not acylating) | AAW01_RS11975 | AAW01_RS00220 |
4-hydroxybenzoate | mhpD: 2-hydroxypentadienoate hydratase | | |
4-hydroxybenzoate | mhpE: 4-hydroxy-2-oxovalerate aldolase | | |
4-hydroxybenzoate | pcaK: 4-hydroxybenzoate transporter pcaK | | |
4-hydroxybenzoate | pobA: 4-hydroxybenzoate 3-monooxygenase | | |
4-hydroxybenzoate | praA: protocatechuate 2,3-dioxygenase | | |
4-hydroxybenzoate | xylF: 2-hydroxymuconate semialdehyde hydrolase | | |
arabinose | araA: L-arabinose isomerase | | |
arabinose | araB: ribulokinase | | |
arabinose | araD: L-ribulose-5-phosphate epimerase | | |
arabinose | araE: L-arabinose:H+ symporter | | |
arginine | astA: arginine N-succinyltransferase | AAW01_RS05075 | |
arginine | astC: succinylornithine transaminase | AAW01_RS09055 | |
arginine | astE: succinylglutamate desuccinylase | | |
arginine | rocE: L-arginine permease | | |
aspartate | glt: aspartate:proton symporter Glt | AAW01_RS13020 | |
cellobiose | glk: glucokinase | AAW01_RS12295 | |
cellobiose | MFS-glucose: glucose transporter, MFS superfamily | AAW01_RS10155 | |
citrate | SLC13A5: citrate:Na+ symporter | | |
citrulline | AO353_03040: ABC transporter for L-Citrulline, ATPase component | AAW01_RS09690 | AAW01_RS11525 |
citrulline | AO353_03045: ABC transporter for L-Citrulline, permease component 2 | | |
citrulline | AO353_03050: ABC transporter for L-Citrulline, permease component 1 | | |
citrulline | AO353_03055: ABC transporter for L-Citrulline, periplasmic substrate-binding component | | |
citrulline | citrullinase: putative citrullinase | AAW01_RS09855 | |
citrulline | ocd: ornithine cyclodeaminase | | |
D-alanine | cycA: D-alanine:H+ symporter CycA | | |
D-alanine | dadA: D-alanine dehydrogenase | | |
D-lactate | D-LDH: D-lactate dehydrogenase | AAW01_RS00865 | AAW01_RS11705 |
D-lactate | lctP: D-lactate:H+ symporter LctP or LidP | | |
D-serine | dsdA: D-serine ammonia-lyase | AAW01_RS06070 | AAW01_RS03555 |
D-serine | dsdX: D-serine transporter DsdX | AAW01_RS06720 | |
deoxyinosine | adh: acetaldehyde dehydrogenase (not acylating) | AAW01_RS11975 | AAW01_RS00220 |
deoxyinosine | deoB: phosphopentomutase | AAW01_RS12560 | |
deoxyinosine | deoC: deoxyribose-5-phosphate aldolase | | |
deoxyinosine | deoD: deoxyinosine phosphorylase | AAW01_RS06200 | |
deoxyinosine | nupC: deoxyinosine:H+ symporter NupC | AAW01_RS05185 | |
deoxyribonate | deoxyribonate-dehyd: 2-deoxy-D-ribonate 3-dehydrogenase | AAW01_RS06580 | AAW01_RS03665 |
deoxyribonate | deoxyribonate-transport: 2-deoxy-D-ribonate transporter | | |
deoxyribonate | garK: glycerate 2-kinase | | |
deoxyribonate | ketodeoxyribonate-cleavage: 2-deoxy-3-keto-D-ribonate cleavage enzyme | | |
deoxyribose | adh: acetaldehyde dehydrogenase (not acylating) | AAW01_RS11975 | AAW01_RS00220 |
deoxyribose | deoC: deoxyribose-5-phosphate aldolase | | |
deoxyribose | deoK: deoxyribokinase | AAW01_RS08110 | |
deoxyribose | deoP: deoxyribose transporter | | |
ethanol | adh: acetaldehyde dehydrogenase (not acylating) | AAW01_RS11975 | AAW01_RS00220 |
fructose | 1pfk: 1-phosphofructokinase | | |
fructose | fruII-ABC: fructose-specific PTS system (fructose 1-phosphate forming), EII-ABC components | | |
fucose | aldA: lactaldehyde dehydrogenase | AAW01_RS04300 | AAW01_RS09770 |
fucose | fucA: L-fuculose-phosphate aldolase FucA | | |
fucose | fucI: L-fucose isomerase FucI | | |
fucose | fucK: L-fuculose kinase FucK | | |
fucose | fucP: L-fucose:H+ symporter FucP | AAW01_RS10155 | |
fucose | fucU: L-fucose mutarotase FucU | | |
galactose | galE: UDP-glucose 4-epimerase | AAW01_RS13100 | AAW01_RS10190 |
galactose | galK: galactokinase (-1-phosphate forming) | | |
galactose | galT: UDP-glucose:alpha-D-galactose-1-phosphate uridylyltransferase | | |
galactose | HP1174: Na+-dependent galactose transporter | AAW01_RS10155 | |
galacturonate | exuT: D-galacturonate transporter ExuT | | |
galacturonate | uxaA: D-altronate dehydratase | AAW01_RS05290 | |
galacturonate | uxaB: tagaturonate reductase | | |
galacturonate | uxaC: D-galacturonate isomerase | AAW01_RS05315 | |
gluconate | gntK: D-gluconate kinase | | |
gluconate | gntT: gluconate:H+ symporter GntT | AAW01_RS06720 | |
glucosamine | gamP: glucosamine PTS system, EII-CBA components (GamP/NagE) | | |
glucosamine | nagB: glucosamine 6-phosphate deaminase (isomerizing) | AAW01_RS04965 | |
glucose | glk: glucokinase | AAW01_RS12295 | |
glucose | MFS-glucose: glucose transporter, MFS superfamily | AAW01_RS10155 | |
glucose-6-P | uhpT: glucose-6-phosphate:phosphate antiporter | | |
glucuronate | dctQ: D-glucuronate TRAP transporter, small permease component | AAW01_RS05305 | |
glucuronate | uxaC: D-glucuronate isomerase | AAW01_RS05315 | |
glutamate | gdhA: glutamate dehydrogenase, NAD-dependent | AAW01_RS07660 | AAW01_RS13210 |
glycerol | glpD: glycerol 3-phosphate dehydrogenase (monomeric) | AAW01_RS13670 | |
glycerol | glpF: glycerol facilitator glpF | AAW01_RS09185 | |
histidine | hutG: N-formiminoglutamate formiminohydrolase | | |
histidine | hutH: histidine ammonia-lyase | | |
histidine | hutI: imidazole-5-propionate hydrolase | | |
histidine | hutU: urocanase | | |
histidine | permease: L-histidine permease | | |
isoleucine | acdH: (2S)-2-methylbutanoyl-CoA dehydrogenase | AAW01_RS10540 | AAW01_RS07145 |
isoleucine | Bap2: L-isoleucine permease Bap2 | | |
L-malate | sdlC: L-malate:Na+ symporter SdlC | | |
lactose | galE: UDP-glucose 4-epimerase | AAW01_RS13100 | AAW01_RS10190 |
lactose | galK: galactokinase (-1-phosphate forming) | | |
lactose | galT: UDP-glucose:alpha-D-galactose-1-phosphate uridylyltransferase | | |
lactose | glk: glucokinase | AAW01_RS12295 | |
lactose | lacP: lactose permease LacP | | |
lactose | lacZ: lactase (homomeric) | | |
leucine | leuT: L-leucine:Na+ symporter LeuT | AAW01_RS01765 | |
leucine | liuC: 3-methylglutaconyl-CoA hydratase | AAW01_RS13205 | AAW01_RS11465 |
leucine | liuD: 3-methylcrotonyl-CoA carboxylase, beta subunit | AAW01_RS07130 | AAW01_RS04080 |
leucine | liuE: hydroxymethylglutaryl-CoA lyase | AAW01_RS11300 | |
lysine | davA: 5-aminovaleramidase | AAW01_RS00785 | AAW01_RS09855 |
lysine | davB: L-lysine 2-monooxygenase | | |
lysine | davD: glutarate semialdehyde dehydrogenase | AAW01_RS09770 | AAW01_RS04300 |
lysine | davT: 5-aminovalerate aminotransferase | AAW01_RS09055 | AAW01_RS12605 |
lysine | fadB: (S)-3-hydroxybutanoyl-CoA dehydrogenase | AAW01_RS00825 | AAW01_RS06205 |
lysine | gcdG: succinyl-CoA:glutarate CoA-transferase | AAW01_RS11415 | AAW01_RS11420 |
lysine | lysP: L-lysine:H+ symporter LysP | | |
maltose | glk: glucokinase | AAW01_RS12295 | |
mannitol | mtlA: mannitol phosphotransferase system, EII-CBA components | | |
mannitol | mtlD: mannitol-1-phosphate 5-dehydrogenase | | |
mannose | manA: mannose-6-phosphate isomerase | AAW01_RS09890 | AAW01_RS13200 |
mannose | manP: mannose PTS system, EII-CBA components | | |
myoinositol | iolG: myo-inositol 2-dehydrogenase | | |
myoinositol | iolM: 2-inosose 4-dehydrogenase | | |
myoinositol | iolN: 2,4-diketo-inositol hydratase | | |
myoinositol | iolO: 5-dehydro-L-gluconate epimerase | | |
myoinositol | iolT: myo-inositol:H+ symporter | | |
myoinositol | uxaE: D-tagaturonate epimerase | | |
NAG | nagA: N-acetylglucosamine 6-phosphate deacetylase | | |
NAG | nagB: glucosamine 6-phosphate deaminase (isomerizing) | AAW01_RS04965 | |
NAG | nagEcba: N-acetylglucosamine phosphotransferase system, EII-CBA components | | |
phenylacetate | paaA: phenylacetyl-CoA 1,2-epoxidase, subunit A | | |
phenylacetate | paaB: phenylacetyl-CoA 1,2-epoxidase, subunit B | | |
phenylacetate | paaC: phenylacetyl-CoA 1,2-epoxidase, subunit C | | |
phenylacetate | paaE: phenylacetyl-CoA 1,2-epoxidase, subunit E | | |
phenylacetate | paaF: 2,3-dehydroadipyl-CoA hydratase | AAW01_RS13205 | AAW01_RS03435 |
phenylacetate | paaH: 3-hydroxyadipyl-CoA dehydrogenase | AAW01_RS00825 | AAW01_RS06205 |
phenylacetate | paaJ1: 3-oxo-5,6-dehydrosuberyl-CoA thiolase | AAW01_RS00830 | AAW01_RS02970 |
phenylacetate | paaJ2: 3-oxoadipyl-CoA thiolase | AAW01_RS00830 | AAW01_RS02970 |
phenylacetate | paaK: phenylacetate-CoA ligase | AAW01_RS11440 | AAW01_RS00835 |
phenylacetate | paaT: phenylacetate transporter Paa | | |
phenylacetate | paaZ1: oxepin-CoA hydrolase | AAW01_RS13205 | |
phenylacetate | paaZ2: 3-oxo-5,6-didehydrosuberyl-CoA semialdehyde dehydrogenase | | |
phenylalanine | aroP: L-phenylalanine:H+ symporter AroP | | |
phenylalanine | QDPR: 6,7-dihydropteridine reductase | | |
propionate | putP: propionate transporter; proline:Na+ symporter | AAW01_RS00770 | |
putrescine | gabT: gamma-aminobutyrate transaminase | AAW01_RS09055 | AAW01_RS12605 |
putrescine | patA: putrescine aminotransferase (PatA/SpuC) | AAW01_RS09055 | AAW01_RS12605 |
putrescine | patD: gamma-aminobutyraldehyde dehydrogenase | AAW01_RS09770 | AAW01_RS04390 |
putrescine | puuP: putrescine:H+ symporter PuuP/PlaP | | |
pyruvate | actP: large subunit of pyruvate transporter (actP-like) | AAW01_RS03875 | |
pyruvate | yjcH: putative small subunit of pyruvate transporter (yjcH-like) | AAW01_RS03880 | |
rhamnose | aldA: lactaldehyde dehydrogenase | AAW01_RS04300 | AAW01_RS09770 |
rhamnose | rhaA: L-rhamnose isomerase | | |
rhamnose | rhaB: L-rhamnulokinase | | |
rhamnose | rhaD: rhamnulose 1-phosphate aldolase | | |
rhamnose | rhaM: L-rhamnose mutarotase | | |
rhamnose | rhaT: L-rhamnose:H+ symporter RhaT | | |
ribose | rbsK: ribokinase | AAW01_RS08110 | |
ribose | rbsU: probable D-ribose transporter RbsU | | |
serine | snatA: L-serine transporter | AAW01_RS06870 | |
sorbitol | mtlA: PTS system for polyols, EII-CBA components | | |
sorbitol | srlD: sorbitol 6-phosphate 2-dehydrogenase | AAW01_RS11455 | AAW01_RS01220 |
succinate | sdc: succinate:Na+ symporter Sdc | | |
sucrose | ams: sucrose hydrolase (invertase) | AAW01_RS05140 | |
sucrose | glk: glucokinase | AAW01_RS12295 | |
sucrose | MFS-glucose: glucose transporter, MFS superfamily | AAW01_RS10155 | |
threonine | gcvP: glycine cleavage system, P component (glycine decarboxylase) | AAW01_RS10165 | AAW01_RS10170 |
threonine | snatA: L-threonine transporter snatA | AAW01_RS06870 | |
thymidine | adh: acetaldehyde dehydrogenase (not acylating) | AAW01_RS11975 | AAW01_RS00220 |
thymidine | deoA: thymidine phosphorylase DeoA | | |
thymidine | deoB: phosphopentomutase | AAW01_RS12560 | |
thymidine | deoC: deoxyribose-5-phosphate aldolase | | |
thymidine | nupG: thymidine permease NupG/XapB | | |
trehalose | glk: glucokinase | AAW01_RS12295 | |
trehalose | MFS-glucose: glucose transporter, MFS superfamily | AAW01_RS10155 | |
trehalose | treF: trehalase | AAW01_RS05140 | AAW01_RS03580 |
tryptophan | aroP: tryptophan:H+ symporter AroP | | |
tryptophan | tnaA: tryptophanase | | |
tyrosine | aroP: L-tyrosine transporter (AroP/FywP) | | |
valine | acdH: isobutyryl-CoA dehydrogenase | AAW01_RS10540 | AAW01_RS07145 |
valine | Bap2: L-valine permease Bap2 | | |
xylitol | fruI: xylitol PTS, enzyme IIABC (FruI) | | |
xylitol | x5p-reductase: D-xylulose-5-phosphate 2-reductase | AAW01_RS09765 | |
xylose | xylA: xylose isomerase | | |
xylose | xylB: xylulokinase | | |
xylose | xylT: D-xylose transporter | | |
Confidence: high confidence medium confidence low confidence
This GapMind analysis is from Sep 24 2021. The underlying query database was built on Sep 17 2021.
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About GapMind
Each pathway is defined by a set of rules based on individual steps or genes. Candidates for each step are identified by using
ublast (a fast alternative to protein BLAST)
against a database of manually-curated proteins (most of which are experimentally characterized) or by using
HMMer with enzyme models (usually from
TIGRFam). Ublast hits may be split across two different proteins.
A candidate for a step is "high confidence" if either:
- ublast finds a hit to a characterized protein at above 40% identity and 80% coverage, and bits >= other bits+10.
- (Hits to curated proteins without experimental data as to their function are never considered high confidence.)
- HMMer finds a hit with 80% coverage of the model, and either other identity < 40 or other coverage < 0.75.
where "other" refers to the best ublast hit to a sequence that is not annotated as performing this step (and is not "ignored").
Otherwise, a candidate is "medium confidence" if either:
- ublast finds a hit at above 40% identity and 70% coverage (ignoring otherBits).
- ublast finds a hit at above 30% identity and 80% coverage, and bits >= other bits.
- HMMer finds a hit (regardless of coverage or other bits).
Other blast hits with at least 50% coverage are "low confidence."
Steps with no high- or medium-confidence candidates may be considered "gaps."
For the typical bacterium that can make all 20 amino acids, there are 1-2 gaps in amino acid biosynthesis pathways.
For diverse bacteria and archaea that can utilize a carbon source, there is a complete
high-confidence catabolic pathway (including a transporter) just 38% of the time, and
there is a complete medium-confidence pathway 63% of the time.
Gaps may be due to:
- our ignorance of proteins' functions,
- omissions in the gene models,
- frame-shift errors in the genome sequence, or
- the organism lacks the pathway.
GapMind relies on the predicted proteins in the genome and does not search the six-frame translation. In most cases, you can search the six-frame translation by clicking on links to Curated BLAST for each step definition (in the per-step page).
For more information, see:
If you notice any errors or omissions in the step descriptions, or any questionable results, please let us know
by Morgan Price, Arkin group, Lawrence Berkeley National Laboratory