PaperBLAST – Find papers about a protein or its homologs

 

PaperBLAST

PaperBLAST Hits for SwissProt::P30145 Sodium/proton-dependent alanine carrier protein (Bacillus sp. (strain PS3)) (445 a.a., MIRLVTMGKS...)

Other sequence analysis tools:

Find functional residues: SitesBLAST

Search for conserved domains

Find the best match in UniProt

Compare to protein structures

Predict transmenbrane helices: Phobius

Predict protein localization: PSORTb

Find homologs in fast.genomics

Fitness BLAST: loading...

Found 126 similar proteins in the literature:

ALCP_BACP3 / P30145 Sodium/proton-dependent alanine carrier protein from Bacillus sp. (strain PS3) (see paper)
TC 2.A.25.1.2 / P30145 Alanine:Na+ (or H+) symporter (see paper)
100% identity, 100% coverage

BAS0605 sodium/alanine symporter family protein from Bacillus anthracis str. Sterne
57% identity, 91% coverage

NP_390659 sodium/proton-dependent alanine transporter from Bacillus subtilis subsp. subtilis str. 168
60% identity, 91% coverage

SO3063 sodium:alanine symporter family protein from Shewanella oneidensis MR-1
55% identity, 87% coverage

BAS2639 sodium/alanine symporter family protein from Bacillus anthracis str. Sterne
BA2830 sodium/alanine symporter family protein from Bacillus anthracis str. Ames
52% identity, 89% coverage

HD73_3147 alanine/glycine:cation symporter family protein from Bacillus thuringiensis serovar kurstaki str. HD73
54% identity, 89% coverage

VP0622 alanine/glycine:cation symporter family protein from Vibrio parahaemolyticus RIMD 2210633
51% identity, 86% coverage

A0QRB1 Amino acid carrier protein from Mycolicibacterium smegmatis (strain ATCC 700084 / mc(2)155)
MSMEG_1052 amino acid carrier protein from Mycobacterium smegmatis str. MC2 155
MSMEG_2332 amino acid carrier protein from Mycobacterium smegmatis str. MC2 155
MSMEG_1052, MSMEG_2332 alanine/glycine:cation symporter family protein from Mycolicibacterium smegmatis MC2 155
49% identity, 84% coverage

WP_047131695 alanine/glycine:cation symporter family protein from Staphylococcus condimenti
48% identity, 80% coverage

SXYL_01919 alanine/glycine:cation symporter family protein from Staphylococcus xylosus
47% identity, 84% coverage

VIBR0546_18581 alanine/glycine:cation symporter family protein from Vibrio brasiliensis LMG 20546
50% identity, 87% coverage

VC0784 sodium/alanine symporter from Vibrio cholerae O1 biovar eltor str. N16961
50% identity, 86% coverage

SA0871 hypothetical protein from Staphylococcus aureus subsp. aureus N315
47% identity, 88% coverage

NWMN_0883 Na+/alanine symporter family protein from Staphylococcus aureus subsp. aureus str. Newman
47% identity, 82% coverage

SAUSA300_0914 sodium:alanine symporter family protein from Staphylococcus aureus subsp. aureus USA300_FPR3757
SACOL1018 sodium:alanine symporter family protein from Staphylococcus aureus subsp. aureus COL
47% identity, 85% coverage

SAFDA_0883 alanine/glycine:cation symporter family protein from Staphylococcus aureus
47% identity, 85% coverage

SAOUHSC_00949 hypothetical protein from Staphylococcus aureus subsp. aureus NCTC 8325
47% identity, 85% coverage

GBAA5301 sodium/alanine symporter family protein from Bacillus anthracis str. 'Ames Ancestor'
GBAA_5301 alanine/glycine:cation symporter family protein from Bacillus anthracis str. 'Ames Ancestor'
41% identity, 92% coverage

BCE_5200 sodium/alanine symporter family protein from Bacillus cereus ATCC 10987
39% identity, 90% coverage

VC2356 sodium/alanine symporter from Vibrio cholerae O1 biovar eltor str. N16961
41% identity, 91% coverage

U876_05745 alanine/glycine:cation symporter family protein from Aeromonas hydrophila NJ-35
42% identity, 88% coverage

GLNT_BACSU / O31464 Probable sodium/glutamine symporter GlnT from Bacillus subtilis (strain 168) (see paper)
U712_01235 alanine/glycine:cation symporter family protein from Bacillus subtilis PY79
BSU02420 glutamine transporter from Bacillus subtilis subsp. subtilis str. 168
40% identity, 93% coverage

NGO1787 putative amino-acid transport protein from Neisseria gonorrhoeae FA 1090
40% identity, 88% coverage

NMA0073 putative amino-acid transport protein from Neisseria meningitidis Z2491
40% identity, 88% coverage

HMPREF0389_01064 alanine/glycine:cation symporter family protein from Filifactor alocis ATCC 35896
38% identity, 91% coverage

PMI1601 sodium:alanine symporter from Proteus mirabilis HI4320
39% identity, 85% coverage

GYO_2182 alanine/glycine:cation symporter family protein from Bacillus spizizenii TU-B-10
38% identity, 93% coverage

PP0496 sodium/alanine symporter from Pseudomonas putida KT2440
38% identity, 85% coverage

SE1038 amino acid carrier protein from Staphylococcus epidermidis ATCC 12228
36% identity, 86% coverage

TC 2.A.25.1.5 / Q45068 Amino-acid carrier protein, AlsT from Bacillus subtilis (strain 168) (see 4 papers)
37% identity, 93% coverage

PLES_13941 putative amino acid permease from Pseudomonas aeruginosa LESB58
PA3641 probable amino acid permease from Pseudomonas aeruginosa PAO1
37% identity, 92% coverage

WP_047131152 alanine/glycine:cation symporter family protein from Staphylococcus condimenti
36% identity, 89% coverage

HI0183 amino acid carrier protein, putative from Haemophilus influenzae Rd KW20
36% identity, 89% coverage

FN1398 Amino acid carrier protein alsT from Fusobacterium nucleatum subsp. nucleatum ATCC 25586
35% identity, 90% coverage

PA14_35460 putative sodium/alanine symporter from Pseudomonas aeruginosa UCBPP-PA14
37% identity, 85% coverage

PA2252 probable AGCS sodium/alanine/glycine symporter from Pseudomonas aeruginosa PAO1
37% identity, 85% coverage

SACOL1392 sodium:alanine symporter family protein from Staphylococcus aureus subsp. aureus COL
38% identity, 85% coverage

SATW20_13570 alanine/glycine:cation symporter family protein from Staphylococcus aureus subsp. aureus TW20
SAV1356 amino acid carrier protein from Staphylococcus aureus subsp. aureus Mu50
SAOUHSC_01354 sodium:alanine symporter family protein, putative from Staphylococcus aureus subsp. aureus NCTC 8325
SAUSA300_1252 amino acid carrier protein from Staphylococcus aureus subsp. aureus USA300_FPR3757
NWMN_1269 sodium:alanine symporter family protein from Staphylococcus aureus subsp. aureus str. Newman
38% identity, 86% coverage

BMEII0783 NA(+)-LINKED D-ALANINE GLYCINE PERMEASE from Brucella melitensis 16M
40% identity, 87% coverage

NMB1647 putative amino acid symporter from Neisseria meningitidis MC58
37% identity, 88% coverage

PM0529 unknown from Pasteurella multocida subsp. multocida str. Pm70
35% identity, 86% coverage

PBPRA1278 hypothetical amino acid carrier protein(sodium/alanine symporter) from Photobacterium profundum SS9
34% identity, 90% coverage

CJJ81176_0912 amino acid carrier protein from Campylobacter jejuni subsp. jejuni 81-176
35% identity, 86% coverage

Cj0903c putative amino-acid transport protein from Campylobacter jejuni subsp. jejuni NCTC 11168
CJSA_0849 alanine/glycine:cation symporter family protein from Campylobacter jejuni subsp. jejuni IA3902
36% identity, 85% coverage

NMA1901 putative sodium:alanine symporter from Neisseria meningitidis Z2491
36% identity, 88% coverage

NGO1290 putative amino-acid transporter, sodium/alanine symporter from Neisseria gonorrhoeae FA 1090
36% identity, 88% coverage

SXYL_01528 alanine/glycine:cation symporter family protein from Staphylococcus xylosus
34% identity, 90% coverage

DV527_RS07535 alanine/glycine:cation symporter family protein from Staphylococcus saprophyticus
36% identity, 90% coverage

A1S_2793 putative amino-acid transport protein from Acinetobacter baumannii ATCC 17978
37% identity, 93% coverage

AbA118F_2024 alanine/glycine:cation symporter family protein from Acinetobacter baumannii
ABAYE0689 putative amino-acid transport protein from Acinetobacter baumannii AYE
37% identity, 84% coverage

TDE1259 amino acid carrier family protein from Treponema denticola ATCC 35405
37% identity, 87% coverage

BP951000_1860 alanine/glycine:cation symporter family protein from Brachyspira pilosicoli 95/1000
38% identity, 84% coverage

cg0254 amino acid carrier protein (sodium/alanine symporter) from Corynebacterium glutamicum ATCC 13032
36% identity, 85% coverage

FN0328 Na(+)-linked D-alanine glycine permease from Fusobacterium nucleatum subsp. nucleatum ATCC 25586
37% identity, 84% coverage

TP0998 sodium/proton-dependent alanine transporter from Treponema pallidum subsp. pallidum str. Nichols
34% identity, 81% coverage

STM0006 putative AGCS family, alanine/glycine transport protein from Salmonella typhimurium LT2
33% identity, 86% coverage

t0006 putative amino-acid transport protein from Salmonella enterica subsp. enterica serovar Typhi Ty2
33% identity, 86% coverage

BP951000_1859 alanine/glycine:cation symporter family protein from Brachyspira pilosicoli 95/1000
37% identity, 87% coverage

AGCS_METMP / Q6LX42 Sodium/alanine symporter AgcS; Alanine permease; Alanine/glycine:cation symporter from Methanococcus maripaludis (strain DSM 14266 / JCM 13030 / NBRC 101832 / S2 / LL) (see 2 papers)
TC 2.A.25.1.3 / Q6LX42 Alanine:Na+ symporter, AgcS from Methanococcus maripaludis (see paper)
MMP1511 Sodium:alanine symporter from Methanococcus maripaludis S2
36% identity, 89% coverage

6cseM / Q6LX42 Crystal structure of sodium/alanine symporter agcs with l-alanine bound (see paper)
36% identity, 92% coverage

PH0781 alanine transport protein from Pyrococcus horikoshii OT3
31% identity, 77% coverage

PF0514 d-alanine glycine permease from Pyrococcus furiosus DSM 3638
31% identity, 74% coverage

APL_1127 predicted Na+/alanine symporter from Actinobacillus pleuropneumoniae L20
35% identity, 89% coverage

APJL_1146 Na+/alanine symporter from Actinobacillus pleuropneumoniae serovar 3 str. JL03
35% identity, 85% coverage

HI0883 amino acid carrier protein, putative from Haemophilus influenzae Rd KW20
35% identity, 88% coverage

Z0007 inner membrane transport protein from Escherichia coli O157:H7 EDL933
33% identity, 87% coverage

YaaJ / b0007 putative transporter YaaJ from Escherichia coli K-12 substr. MG1655 (see 4 papers)
P30143 Uncharacterized transporter YaaJ from Escherichia coli (strain K12)
b0007 predicted transporter from Escherichia coli str. K-12 substr. MG1655
33% identity, 87% coverage

NTHI1046 predicted Na+/alanine symporter from Haemophilus influenzae 86-028NP
35% identity, 88% coverage

PMI2114 sodium:alanine symporter from Proteus mirabilis HI4320
35% identity, 88% coverage

DEFDS_1601 sodium:alanine symporter from Deferribacter desulfuricans SSM1
36% identity, 86% coverage

PAP_09570 alanine/glycine:cation symporter family protein from Palaeococcus pacificus DY20341
31% identity, 88% coverage

NMB0177 putative sodium/alanine symporter from Neisseria meningitidis MC58
33% identity, 88% coverage

SAG1145 sodium:alanine symporter family protein from Streptococcus agalactiae 2603V/R
33% identity, 92% coverage

gbs1212 Unknown from Streptococcus agalactiae NEM316
33% identity, 92% coverage

NMA0091 putative amino-acid transporter from Neisseria meningitidis Z2491
33% identity, 88% coverage

NGFG_01721 alanine/glycine:cation symporter family protein from Neisseria gonorrhoeae MS11
NGO1807 putative amino-acid transporter from Neisseria gonorrhoeae FA 1090
33% identity, 88% coverage

SPD_0372 sodium:alanine symporter family protein from Streptococcus pneumoniae D39
spr0369 D-alanine glycine permease from Streptococcus pneumoniae R6
33% identity, 91% coverage

BH4033 amino acid transporter from Bacillus halodurans C-125
32% identity, 88% coverage

NSU_2100 alanine/glycine:cation symporter family protein from Novosphingobium pentaromativorans US6-1
32% identity, 75% coverage

PP1Y_AT4545 alanine/glycine:cation symporter family protein from Novosphingobium sp. PP1Y
32% identity, 75% coverage

STACADC2_0872 alanine/glycine:cation symporter family protein from Streptococcus thermophilus
33% identity, 91% coverage

AWO_RS12525, Awo_c24340 alanine/glycine:cation symporter family protein from Acetobacterium woodii DSM 1030
31% identity, 80% coverage

TC 2.A.25.1.9 / W0WFC6 Probable Glycine/alanine/asparagine/glutamine uptake porter, AgcS from Pseudomonas aeruginosa MH38
PA2533 probable sodium:alanine symporter from Pseudomonas aeruginosa PAO1
31% identity, 90% coverage

Spy49_1008 Putative amino acid symporter from Streptococcus pyogenes NZ131
33% identity, 91% coverage

PM0091 unknown from Pasteurella multocida subsp. multocida str. Pm70
PmVP161_1144 alanine/glycine:cation symporter family protein from Pasteurella multocida
32% identity, 90% coverage

BC2317 Na(+)-linked D-alanine glycine permease from Bacillus cereus ATCC 14579
34% identity, 84% coverage

SPy1270, SPy_1270 putative amino acid symporter from Streptococcus pyogenes M1 GAS
33% identity, 91% coverage

ABO_0618 sodium/alanine symporter from Alcanivorax borkumensis SK2
32% identity, 84% coverage

CD2276 putative sodium:solute symporter from Clostridium difficile 630
33% identity, 88% coverage

Pro1538, SS120_15671 Na+/alanine symporter from Prochlorococcus marinus str. SS120
34% identity, 85% coverage

Pcar_2492 sodium/alanine symporter from Pelobacter carbinolicus str. DSM 2380
33% identity, 85% coverage

VIBR0546_12742 alanine/glycine:cation symporter family protein from Vibrio brasiliensis LMG 20546
31% identity, 87% coverage

HP0942 D-alanine glycine permease (dagA) from Helicobacter pylori 26695
30% identity, 88% coverage

Elen_1016 amino acid carrier protein from Eggerthella lenta DSM 2243
30% identity, 87% coverage

VP1741 sodium/alanine symporter from Vibrio parahaemolyticus RIMD 2210633
30% identity, 89% coverage

BSU07750 putative aminoacid transporter from Bacillus subtilis subsp. subtilis str. 168
36% identity, 61% coverage

SO0858 sodium:alanine symporter family protein from Shewanella oneidensis MR-1
34% identity, 84% coverage

jhp0877 putative SODIUM/ALANINE SYMPORTER from Helicobacter pylori J99
30% identity, 88% coverage

VC1422 sodium/alanine symporter from Vibrio cholerae O1 biovar eltor str. N16961
30% identity, 89% coverage

SPO2370 sodium:alanine symporter family protein from Silicibacter pomeroyi DSS-3
31% identity, 77% coverage

SPO2370 alanine/glycine:cation symporter family protein from Ruegeria pomeroyi DSS-3
31% identity, 80% coverage

SMU_1175 alanine/glycine:cation symporter family protein from Streptococcus mutans UA159
31% identity, 92% coverage

DVU0724 sodium/alanine symporter family protein from Desulfovibrio vulgaris Hildenborough
33% identity, 82% coverage

FN1747 Cysteine permease from Fusobacterium nucleatum subsp. nucleatum ATCC 25586
30% identity, 80% coverage

IL1268 Na(+)-linked D-alanine and glycine permease from Idiomarina loihiensis L2TR
35% identity, 47% coverage

DAGA_PSEHA / P30144 Na(+)-linked D-alanine glycine permease; Sodium/alanine symporter from Pseudoalteromonas haloplanktis (Alteromonas haloplanktis) (see 2 papers)
TC 2.A.25.1.1 / P30144 Alanine (or glycine):Na+ symporter from Alteromonas haloplanktis (see paper)
34% identity, 49% coverage

Q7UKE2 Na(+)-linked D-alanine glycine permease from Rhodopirellula baltica (strain DSM 10527 / NCIMB 13988 / SH1)
31% identity, 72% coverage

CLOSCI_01264 hypothetical protein from Clostridium scindens ATCC 35704
31% identity, 86% coverage

CLOSCI_01264 alanine/glycine:cation symporter family protein from [Clostridium] scindens ATCC 35704
31% identity, 88% coverage

TP0414 D-alanine glycine permease (dagA) from Treponema pallidum subsp. pallidum str. Nichols
31% identity, 88% coverage

pc0622 putative D-alanine/glycine transport protein, sodium-dependent from Parachlamydia sp. UWE25
29% identity, 86% coverage

AMK26_RS14790 alanine/glycine:cation symporter family protein from Streptomyces sp. CB03234
28% identity, 81% coverage

Wbm0424 Na+/alanine symporter from Wolbachia endosymbiont strain TRS of Brugia malayi
26% identity, 88% coverage

CT735 D-Ala/Gly Permease from Chlamydia trachomatis D/UW-3/CX
25% identity, 88% coverage

SEN0907 putative ion:amino acid symporter from Salmonella enterica subsp. enterica serovar Enteritidis str. P125109
26% identity, 85% coverage

CPn0536 D-Ala/Gly Permease from Chlamydophila pneumoniae CWL029
26% identity, 62% coverage

HMPREF0389_01240 alanine/glycine:cation symporter family protein from Filifactor alocis ATCC 35896
31% identity, 53% coverage

CT409 Amino acid permease from Chlamydia trachomatis D/UW-3/CX
24% identity, 83% coverage

pc1598 putative D-alanine/glycine transport protein, sodium-dependent from Parachlamydia sp. UWE25
26% identity, 83% coverage

STM14_1138, STMMW_10141 alanine/glycine:cation symporter family protein from Salmonella enterica subsp. enterica serovar Typhimurium str. 14028S
STM1003 putative transcriptional regulator, Lrp family from Salmonella typhimurium LT2
26% identity, 85% coverage

SEET0819_11930 alanine/glycine:cation symporter family protein from Salmonella enterica subsp. enterica serovar Tennessee str.
27% identity, 81% coverage

Wbm0197 Na+/alanine symporter from Wolbachia endosymbiont strain TRS of Brugia malayi
25% identity, 89% coverage

AM882 hypothetical protein from Anaplasma marginale str. St. Maries
26% identity, 85% coverage

EHF_0383 alanine:cation symporter family protein from Ehrlichia japonica
23% identity, 89% coverage

New Search

For advice on how to use these tools together, see Interactive tools for functional annotation of bacterial genomes.

Statistics

The PaperBLAST database links 793,807 different protein sequences to 1,259,118 scientific articles. Searches against EuropePMC were last performed on March 13 2025.

How It Works

PaperBLAST builds a database of protein sequences that are linked to scientific articles. These links come from automated text searches against the articles in EuropePMC and from manually-curated information from GeneRIF, UniProtKB/Swiss-Prot, BRENDA, CAZy (as made available by dbCAN), BioLiP, CharProtDB, MetaCyc, EcoCyc, TCDB, REBASE, the Fitness Browser, and a subset of the European Nucleotide Archive with the /experiment tag. Given this database and a protein sequence query, PaperBLAST uses protein-protein BLAST to find similar sequences with E < 0.001.

To build the database, we query EuropePMC with locus tags, with RefSeq protein identifiers, and with UniProt accessions. We obtain the locus tags from RefSeq or from MicrobesOnline. We use queries of the form "locus_tag AND genus_name" to try to ensure that the paper is actually discussing that gene. Because EuropePMC indexes most recent biomedical papers, even if they are not open access, some of the links may be to papers that you cannot read or that our computers cannot read. We query each of these identifiers that appears in the open access part of EuropePMC, as well as every locus tag that appears in the 500 most-referenced genomes, so that a gene may appear in the PaperBLAST results even though none of the papers that mention it are open access. We also incorporate text-mined links from EuropePMC that link open access articles to UniProt or RefSeq identifiers. (This yields some additional links because EuropePMC uses different heuristics for their text mining than we do.)

For every article that mentions a locus tag, a RefSeq protein identifier, or a UniProt accession, we try to select one or two snippets of text that refer to the protein. If we cannot get access to the full text, we try to select a snippet from the abstract, but unfortunately, unique identifiers such as locus tags are rarely provided in abstracts.

PaperBLAST also incorporates manually-curated protein functions:

Except for GeneRIF and ENA, the curated entries include a short curated description of the protein's function. For entries from BioLiP, the protein's function may not be known beyond binding to the ligand. Many of these entries also link to articles in PubMed.

For more information see the PaperBLAST paper (mSystems 2017) or the code. You can download PaperBLAST's database here.

Changes to PaperBLAST since the paper was written:

Many of these changes are described in Interactive tools for functional annotation of bacterial genomes.

Secrets

PaperBLAST cannot provide snippets for many of the papers that are published in non-open-access journals. This limitation applies even if the paper is marked as "free" on the publisher's web site and is available in PubmedCentral or EuropePMC. If a journal that you publish in is marked as "secret," please consider publishing elsewhere.

Omissions from the PaperBLAST Database

Many important articles are missing from PaperBLAST, either because the article's full text is not in EuropePMC (as for many older articles), or because the paper does not mention a protein identifier such as a locus tag, or because of PaperBLAST's heuristics. If you notice an article that characterizes a protein's function but is missing from PaperBLAST, please notify the curators at UniProt or add an entry to GeneRIF. Entries in either of these databases will eventually be incorporated into PaperBLAST. Note that to add an entry to UniProt, you will need to find the UniProt identifier for the protein. If the protein is not already in UniProt, you can ask them to create an entry. To add an entry to GeneRIF, you will need an NCBI Gene identifier, but unfortunately many prokaryotic proteins in RefSeq do not have corresponding Gene identifers.

References

PaperBLAST: Text-mining papers for information about homologs.
M. N. Price and A. P. Arkin (2017). mSystems, 10.1128/mSystems.00039-17.

Europe PMC in 2017.
M. Levchenko et al (2017). Nucleic Acids Research, 10.1093/nar/gkx1005.

Gene indexing: characterization and analysis of NLM's GeneRIFs.
J. A. Mitchell et al (2003). AMIA Annu Symp Proc 2003:460-464.

UniProt: the universal protein knowledgebase.
The UniProt Consortium (2016). Nucleic Acids Research, 10.1093/nar/gkw1099.

BRENDA in 2017: new perspectives and new tools in BRENDA.
S. Placzek et al (2017). Nucleic Acids Research, 10.1093/nar/gkw952.

The EcoCyc database: reflecting new knowledge about Escherichia coli K-12.
I. M. Keeseler et al (2016). Nucleic Acids Research, 10.1093/nar/gkw1003.

The MetaCyc database of metabolic pathways and enzymes.
R. Caspi et al (2018). Nucleic Acids Research, 10.1093/nar/gkx935.

CharProtDB: a database of experimentally characterized protein annotations.
R. Madupu et al (2012). Nucleic Acids Research, 10.1093/nar/gkr1133.

The carbohydrate-active enzymes database (CAZy) in 2013.
V. Lombard et al (2014). Nucleic Acids Research, 10.1093/nar/gkt1178.

The Transporter Classification Database (TCDB): recent advances
M. H. Saier, Jr. et al (2016). Nucleic Acids Research, 10.1093/nar/gkv1103.

REBASE - a database for DNA restriction and modification: enzymes, genes and genomes.
R. J. Roberts et al (2015). Nucleic Acids Research, 10.1093/nar/gku1046.

Deep annotation of protein function across diverse bacteria from mutant phenotypes.
M. N. Price et al (2016). bioRxiv, 10.1101/072470.

by Morgan Price, Arkin group
Lawrence Berkeley National Laboratory