PaperBLAST – Find papers about a protein or its homologs

 

PaperBLAST

PaperBLAST Hits for EX31_RS22465 (84 a.a., MKPTIHPAYR...)

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Found 98 similar proteins in the literature:

YPK_3210 ribosomal protein L31 from Yersinia pseudotuberculosis YPIII
YPO3134 putative ribosomal protein from Yersinia pestis CO92
y1048 50S ribosomal protein L31 type B from Yersinia pestis KIM
79% identity, 98% coverage

SMDB11_RS01845 type B 50S ribosomal protein L31 from Serratia marcescens subsp. marcescens Db11
77% identity, 100% coverage

ECA_RS05790 type B 50S ribosomal protein L31 from Pectobacterium atrosepticum SCRI1043
81% identity, 96% coverage

KPNJ2_04207 type B 50S ribosomal protein L31 from Klebsiella pneumoniae 30684/NJST258_2
73% identity, 97% coverage

ESA_RS12970 type B 50S ribosomal protein L31 from Cronobacter sakazakii ATCC BAA-894
73% identity, 98% coverage

YkgM / b0296 50S ribosomal subunit protein L31B from Escherichia coli K-12 substr. MG1655 (see 8 papers)
RL31B_ECOLI / P0A7N1 Large ribosomal subunit protein bL31B; 50S ribosomal protein L31 type B from Escherichia coli (strain K12) (see 3 papers)
ECOLIN_01860 type B 50S ribosomal protein L31 from Escherichia coli Nissle 1917
NP_414830 50S ribosomal subunit protein L31B from Escherichia coli str. K-12 substr. MG1655
b0296 50S ribosomal protein L31 from Escherichia coli str. K-12 substr. MG1655
69% identity, 97% coverage

B7L427 Large ribosomal subunit protein bL31B from Escherichia coli (strain 55989 / EAEC)
69% identity, 97% coverage

STM0469 putative 50S ribosomal protein L31 (second copy) from Salmonella typhimurium LT2
SEN0450 putative 50s ribosomal protein L31 (second copy) from Salmonella enterica subsp. enterica serovar Enteritidis str. P125109
JJB80_02375 type B 50S ribosomal protein L31 from Salmonella enterica subsp. enterica serovar Typhimurium
69% identity, 98% coverage

VC0878 ribosomal protein, L31P family from Vibrio cholerae O1 biovar eltor str. N16961
69% identity, 94% coverage

VP_RS11315 type B 50S ribosomal protein L31 from Vibrio parahaemolyticus RIMD 2210633
VP2331 ribosomal protein, L31P family from Vibrio parahaemolyticus RIMD 2210633
65% identity, 98% coverage

U876_13185 type B 50S ribosomal protein L31 from Aeromonas hydrophila NJ-35
63% identity, 93% coverage

PSPTO_4183 ribosomal protein L31 from Pseudomonas syringae pv. tomato str. DC3000
63% identity, 86% coverage

B7C60_RS12925 type B 50S ribosomal protein L31 from Vibrio fujianensis
65% identity, 95% coverage

Psyr_3920 Ribosomal protein L31 from Pseudomonas syringae pv. syringae B728a
60% identity, 90% coverage

6i7vD7 / P0A7N1 6i7vD7 (see paper)
72% identity, 81% coverage

PA14_17700 putative ribosomal protein, L31P family from Pseudomonas aeruginosa UCBPP-PA14
PA3601 hypothetical protein from Pseudomonas aeruginosa PAO1
IS492_27315 type B 50S ribosomal protein L31 from Pseudomonas aeruginosa
59% identity, 97% coverage

PFL_4896 ribosomal protein L31 from Pseudomonas fluorescens Pf-5
60% identity, 90% coverage

HMPREF0010_01851 type B 50S ribosomal protein L31 from Acinetobacter baumannii ATCC 19606 = CIP 70.34 = JCM 6841
A1S_0391 50S ribosomal protein L31 type B from Acinetobacter baumannii ATCC 17978
ABBFA_003147 ribosomal protein L31 from Acinetobacter baumannii AB307-0294
55% identity, 95% coverage

Rmet_2137 type B 50S ribosomal protein L31 from Cupriavidus metallidurans CH34
57% identity, 95% coverage

gbs0581 50S ribosomal protein L31 from Streptococcus agalactiae NEM316
54% identity, 91% coverage

SMU_1298 type B 50S ribosomal protein L31 from Streptococcus mutans UA159
53% identity, 93% coverage

SSA_1272 50S ribosomal protein L31 type B, putative from Streptococcus sanguinis SK36
51% identity, 93% coverage

SPD_1154 ribosomal protein L31 from Streptococcus pneumoniae D39
SP_1299 50S ribosomal protein L31 type B from Streptococcus pneumoniae TIGR4
51% identity, 93% coverage

STER_0787, STER_RS03860 type B 50S ribosomal protein L31 from Streptococcus thermophilus
51% identity, 93% coverage

bglu_1g14290 Ribosomal protein L31 from Burkholderia glumae BGR1
53% identity, 92% coverage

lmo2548 ribosomal protein L31 from Listeria monocytogenes EGD-e
Q71WN0 Large ribosomal subunit protein bL31B from Listeria monocytogenes serotype 4b (strain F2365)
lin2692 ribosomal protein L31 from Listeria innocua Clip11262
53% identity, 93% coverage

7nhn4 / P0A485 7nhn4 (see paper)
53% identity, 93% coverage

BTH_I2213 50S ribosomal protein L31 type B from Burkholderia thailandensis E264
52% identity, 91% coverage

NMB0942 50S ribosomal protein L31 from Neisseria meningitidis MC58
48% identity, 89% coverage

BAS5180 ribosomal protein L31 from Bacillus anthracis str. Sterne
BA5574 ribosomal protein L31 from Bacillus anthracis str. Ames
48% identity, 93% coverage

NMA1138 putative additional 50S ribosomal protein L31 from Neisseria meningitidis Z2491
48% identity, 89% coverage

BPSL1491 50S ribosomal protein L31 from Burkholderia pseudomallei K96243
52% identity, 91% coverage

LLNZ_04655 type B 50S ribosomal protein L31 from Lactococcus cremoris subsp. cremoris NZ9000
49% identity, 93% coverage

5myjB3 / A2RJP7 of 70S ribosome from Lactococcus lactis (see paper)
49% identity, 93% coverage

B4RL59 Large ribosomal subunit protein bL31B from Neisseria gonorrhoeae (strain NCCP11945)
46% identity, 89% coverage

SCO3427 50S ribosomal protein L31 from Streptomyces coelicolor A3(2)
47% identity, 93% coverage

SERP1727 ribosomal protein L31 from Staphylococcus epidermidis RP62A
48% identity, 92% coverage

AS87_RS03960, B739_1112 type B 50S ribosomal protein L31 from Riemerella anatipestifer Yb2
49% identity, 95% coverage

PD0749 50S ribosomal protein L31 from Xylella fastidiosa Temecula1
47% identity, 93% coverage

RL31B_LACPL / Q88Z52 Large ribosomal subunit protein bL31B; 50S ribosomal protein L31 type B from Lactiplantibacillus plantarum (strain ATCC BAA-793 / NCIMB 8826 / WCFS1) (Lactobacillus plantarum) (see paper)
lp_0512 type B 50S ribosomal protein L31 from Lactiplantibacillus plantarum WCFS1
lp_0512 ribosomal protein L31 from Lactobacillus plantarum WCFS1
47% identity, 93% coverage

F452_RS0105140 type B 50S ribosomal protein L31 from Porphyromonas gulae DSM 15663
PGN_0636 50S ribosomal protein L31 from Porphyromonas gingivalis ATCC 33277
PG0592 ribosomal protein L31 from Porphyromonas gingivalis W83
45% identity, 99% coverage

SCO1150 50S ribosomal protein L31 from Streptomyces coelicolor A3(2)
45% identity, 92% coverage

EF1171 ribosomal protein L31 from Enterococcus faecalis V583
IUJ47_RS08865 type B 50S ribosomal protein L31 from Enterococcus faecalis
47% identity, 88% coverage

6o8z1 / A0A1B4XMV6 6o8z1 (see paper)
47% identity, 93% coverage

SA1922 ribosomal protein L31 from Staphylococcus aureus subsp. aureus N315
Q2FF08 Large ribosomal subunit protein bL31B from Staphylococcus aureus (strain USA300)
Q2FWD8 Large ribosomal subunit protein bL31B from Staphylococcus aureus (strain NCTC 8325 / PS 47)
Q6GEV5 Large ribosomal subunit protein bL31B from Staphylococcus aureus (strain MRSA252)
46% identity, 93% coverage

5li03 / Q2FWD8 5li03 (see paper)
46% identity, 93% coverage

cg0994 50S ribosomal protein L31 type B from Corynebacterium glutamicum ATCC 13032
Q8NS12 Large ribosomal subunit protein bL31B from Corynebacterium glutamicum (strain ATCC 13032 / DSM 20300 / JCM 1318 / BCRC 11384 / CCUG 27702 / LMG 3730 / NBRC 12168 / NCIMB 10025 / NRRL B-2784 / 534)
42% identity, 95% coverage

C9Z0W5 Large ribosomal subunit protein bL31B from Streptomyces scabiei (strain 87.22)
SCAB_39991 putative 50S ribosomal protein L31 from Streptomyces scabiei 87.22
44% identity, 99% coverage

LSEI_2569 Ribosomal protein L31 from Lactobacillus casei ATCC 334
46% identity, 93% coverage

E2P69_RS10215, XAC29_17265 type B 50S ribosomal protein L31 from Xanthomonas axonopodis Xac29-1
42% identity, 93% coverage

PMCN03_1019 type B 50S ribosomal protein L31 from Pasteurella multocida subsp. multocida str. HB03
45% identity, 91% coverage

APL_1821 50S ribosomal protein L31 from Actinobacillus pleuropneumoniae L20
43% identity, 92% coverage

CTL0277 LSU ribosomal protein L31P from Chlamydia trachomatis 434/Bu
43% identity, 71% coverage

MAP_RS19325 type B 50S ribosomal protein L31 from Mycobacterium avium subsp. paratuberculosis K-10
MAP3771 hypothetical protein from Mycobacterium avium subsp. paratuberculosis str. k10
41% identity, 81% coverage

MAB_0336 50S ribosomal protein L31 type B from Mycobacterium abscessus ATCC 19977
43% identity, 94% coverage

BB0229 ribosomal protein L31 (rpmE) from Borrelia burgdorferi B31
40% identity, 92% coverage

8uu93 / A0A0U4MT39 8uu93 (see paper)
49% identity, 92% coverage

FN0482 LSU ribosomal protein L31P from Fusobacterium nucleatum subsp. nucleatum ATCC 25586
40% identity, 100% coverage

RL31B_BACSU / O34967 Large ribosomal subunit protein bL31B; 50S ribosomal protein L31 type B from Bacillus subtilis (strain 168) (see paper)
NP_390948 ribosomal protein L31 from Bacillus subtilis subsp. subtilis str. 168
43% identity, 99% coverage

8rd8fJ / A0A0M4SX87 8rd8fJ (see paper)
53% identity, 65% coverage

FP1135 50S ribosomal protein L31 type B from Flavobacterium psychrophilum JIP02/86
IA01_05395 type B 50S ribosomal protein L31 from Flavobacterium psychrophilum
41% identity, 93% coverage

FE46_RS09965 type B 50S ribosomal protein L31 from Flavobacterium psychrophilum
41% identity, 93% coverage

AMUC_RS07650 type B 50S ribosomal protein L31 from Akkermansia muciniphila ATCC BAA-835
37% identity, 87% coverage

CT022 L31 Ribosomal Protein from Chlamydia trachomatis D/UW-3/CX
44% identity, 71% coverage

7p48Y / A0A380EL30 7p48Y (see paper)
41% identity, 93% coverage

MSMEG_6070 ribosomal protein L31 from Mycobacterium smegmatis str. MC2 155
40% identity, 93% coverage

SO4120, SO_4120 ribosomal protein L31 from Shewanella oneidensis MR-1
38% identity, 100% coverage

A0J47_RS02550, VDA_003390 50S ribosomal protein L31 from Photobacterium damselae subsp. damselae
36% identity, 99% coverage

7nhl4 / Q2FWD8 7nhl4 (see paper)
39% identity, 90% coverage

y0299 50S ribosomal protein L31 from Yersinia pestis KIM
YPO0111 50S ribosomal protein L31 from Yersinia pestis CO92
YPTB0102 50S ribosomal protein L31 from Yersinia pseudotuberculosis IP 32953
34% identity, 92% coverage

8cd1Le / Q9HUD0 8cd1Le (see paper)
35% identity, 95% coverage

Q9HUD0 Large ribosomal subunit protein bL31 from Pseudomonas aeruginosa (strain ATCC 15692 / DSM 22644 / CIP 104116 / JCM 14847 / LMG 12228 / 1C / PRS 101 / PAO1)
PA14_66710 50S ribosomal protein L31 from Pseudomonas aeruginosa UCBPP-PA14
PA5049 50S ribosomal protein L31 from Pseudomonas aeruginosa PAO1
35% identity, 95% coverage

PFL_0441 ribosomal protein L31 from Pseudomonas fluorescens Pf-5
32% identity, 95% coverage

PP_5087 ribosomal protein L31 from Pseudomonas putida KT2440
36% identity, 100% coverage

RpmE / b3936 50S ribosomal subunit protein L31 from Escherichia coli K-12 substr. MG1655 (see 22 papers)
rpmE / P0A7M9 50S ribosomal subunit protein L31 from Escherichia coli (strain K12) (see 26 papers)
RL31_ECOLI / P0A7M9 Large ribosomal subunit protein bL31; 50S ribosomal protein L31 from Escherichia coli (strain K12) (see 8 papers)
5akaZ / P0A7M9 Em structure of ribosome-srp-ftsy complex in closed state (see paper)
b3936 50S ribosomal subunit protein L31 from Escherichia coli str. K-12 substr. MG1655
NP_418371 50S ribosomal subunit protein L31 from Escherichia coli str. K-12 substr. MG1655
Z5484 50S ribosomal subunit protein L31 from Escherichia coli O157:H7 EDL933
Z_RS25730 50S ribosomal protein L31 from Escherichia coli O157:H7 str. EDL933
32% identity, 92% coverage

NP_709740 50S ribosomal subunit protein L31 from Shigella flexneri 2a str. 301
32% identity, 92% coverage

VC2679 ribosomal protein L31 from Vibrio cholerae O1 biovar eltor str. N16961
33% identity, 100% coverage

ACIAD2210 50S ribosomal protein L31 from Acinetobacter sp. ADP1
36% identity, 92% coverage

KF946_12560 50S ribosomal protein L31 from Idiomarina loihiensis
37% identity, 100% coverage

VP0255 ribosomal protein L31 from Vibrio parahaemolyticus RIMD 2210633
34% identity, 95% coverage

EAMY_0136 50S ribosomal protein L31 from Erwinia amylovora CFBP1430
32% identity, 92% coverage

HSM_0533 50S ribosomal protein L31 from Histophilus somni 2336
HSM_0533 ribosomal protein L31 from Haemophilus somnus 2336
32% identity, 92% coverage

Asuc_0525 ribosomal protein L31 from Actinobacillus succinogenes 130Z
34% identity, 92% coverage

MT1337 50S ribosomal protein L31 from Mycobacterium tuberculosis CDC1551
Rv1298 50S ribosomal protein L31 from Mycobacterium tuberculosis H37Rv
40% identity, 93% coverage

B7C60_RS02450 50S ribosomal protein L31 from Vibrio fujianensis
35% identity, 100% coverage

MAB_1441 50S ribosomal protein L31 from Mycobacterium abscessus ATCC 19977
37% identity, 93% coverage

7jila / A0A1M5KIG2 7jila (see paper)
43% identity, 68% coverage

HI0758 ribosomal protein L31 (rpL31) from Haemophilus influenzae Rd KW20
34% identity, 92% coverage

APL_0982 50S ribosomal protein L31 from Actinobacillus pleuropneumoniae L20
31% identity, 92% coverage

RL31_STRCO / Q9K4E5 Large ribosomal subunit protein bL31; 50S ribosomal protein L31 from Streptomyces coelicolor (strain ATCC BAA-471 / A3(2) / M145) (see paper)
SCO5359 50S ribosomal protein L31 from Streptomyces coelicolor A3(2)
40% identity, 93% coverage

SYNW2095 50S ribosomal protein L31 from Synechococcus sp. WH 8102
30% identity, 97% coverage

9c4g4 / A0A2B7JUW7 Cutibacterium acnes 50s ribosomal subunit with clindamycin bound (see paper)
35% identity, 93% coverage

5zeb2 / A0R215 5zeb2 (see paper)
38% identity, 93% coverage

MSMEG_4951 50S ribosomal protein L31 from Mycolicibacterium smegmatis MC2 155
MSMEG_4951 ribosomal protein L31 from Mycobacterium smegmatis str. MC2 155
38% identity, 93% coverage

PMT1760 50S ribosomal protein L31 from Prochlorococcus marinus str. MIT 9313
29% identity, 83% coverage

ssl3445 50S ribosomal protein L31 from Synechocystis sp. PCC 6803
45% identity, 48% coverage

SRIM_028075 50S ribosomal protein L31 from Streptomyces rimosus subsp. rimosus ATCC 10970
37% identity, 98% coverage

7nhk3 / A0A1B4XMV6 7nhk3 (see paper)
35% identity, 93% coverage

New Search

For advice on how to use these tools together, see Interactive tools for functional annotation of bacterial genomes.

Statistics

The PaperBLAST database links 793,807 different protein sequences to 1,259,118 scientific articles. Searches against EuropePMC were last performed on March 13 2025.

How It Works

PaperBLAST builds a database of protein sequences that are linked to scientific articles. These links come from automated text searches against the articles in EuropePMC and from manually-curated information from GeneRIF, UniProtKB/Swiss-Prot, BRENDA, CAZy (as made available by dbCAN), BioLiP, CharProtDB, MetaCyc, EcoCyc, TCDB, REBASE, the Fitness Browser, and a subset of the European Nucleotide Archive with the /experiment tag. Given this database and a protein sequence query, PaperBLAST uses protein-protein BLAST to find similar sequences with E < 0.001.

To build the database, we query EuropePMC with locus tags, with RefSeq protein identifiers, and with UniProt accessions. We obtain the locus tags from RefSeq or from MicrobesOnline. We use queries of the form "locus_tag AND genus_name" to try to ensure that the paper is actually discussing that gene. Because EuropePMC indexes most recent biomedical papers, even if they are not open access, some of the links may be to papers that you cannot read or that our computers cannot read. We query each of these identifiers that appears in the open access part of EuropePMC, as well as every locus tag that appears in the 500 most-referenced genomes, so that a gene may appear in the PaperBLAST results even though none of the papers that mention it are open access. We also incorporate text-mined links from EuropePMC that link open access articles to UniProt or RefSeq identifiers. (This yields some additional links because EuropePMC uses different heuristics for their text mining than we do.)

For every article that mentions a locus tag, a RefSeq protein identifier, or a UniProt accession, we try to select one or two snippets of text that refer to the protein. If we cannot get access to the full text, we try to select a snippet from the abstract, but unfortunately, unique identifiers such as locus tags are rarely provided in abstracts.

PaperBLAST also incorporates manually-curated protein functions:

Except for GeneRIF and ENA, the curated entries include a short curated description of the protein's function. For entries from BioLiP, the protein's function may not be known beyond binding to the ligand. Many of these entries also link to articles in PubMed.

For more information see the PaperBLAST paper (mSystems 2017) or the code. You can download PaperBLAST's database here.

Changes to PaperBLAST since the paper was written:

Many of these changes are described in Interactive tools for functional annotation of bacterial genomes.

Secrets

PaperBLAST cannot provide snippets for many of the papers that are published in non-open-access journals. This limitation applies even if the paper is marked as "free" on the publisher's web site and is available in PubmedCentral or EuropePMC. If a journal that you publish in is marked as "secret," please consider publishing elsewhere.

Omissions from the PaperBLAST Database

Many important articles are missing from PaperBLAST, either because the article's full text is not in EuropePMC (as for many older articles), or because the paper does not mention a protein identifier such as a locus tag, or because of PaperBLAST's heuristics. If you notice an article that characterizes a protein's function but is missing from PaperBLAST, please notify the curators at UniProt or add an entry to GeneRIF. Entries in either of these databases will eventually be incorporated into PaperBLAST. Note that to add an entry to UniProt, you will need to find the UniProt identifier for the protein. If the protein is not already in UniProt, you can ask them to create an entry. To add an entry to GeneRIF, you will need an NCBI Gene identifier, but unfortunately many prokaryotic proteins in RefSeq do not have corresponding Gene identifers.

References

PaperBLAST: Text-mining papers for information about homologs.
M. N. Price and A. P. Arkin (2017). mSystems, 10.1128/mSystems.00039-17.

Europe PMC in 2017.
M. Levchenko et al (2017). Nucleic Acids Research, 10.1093/nar/gkx1005.

Gene indexing: characterization and analysis of NLM's GeneRIFs.
J. A. Mitchell et al (2003). AMIA Annu Symp Proc 2003:460-464.

UniProt: the universal protein knowledgebase.
The UniProt Consortium (2016). Nucleic Acids Research, 10.1093/nar/gkw1099.

BRENDA in 2017: new perspectives and new tools in BRENDA.
S. Placzek et al (2017). Nucleic Acids Research, 10.1093/nar/gkw952.

The EcoCyc database: reflecting new knowledge about Escherichia coli K-12.
I. M. Keeseler et al (2016). Nucleic Acids Research, 10.1093/nar/gkw1003.

The MetaCyc database of metabolic pathways and enzymes.
R. Caspi et al (2018). Nucleic Acids Research, 10.1093/nar/gkx935.

CharProtDB: a database of experimentally characterized protein annotations.
R. Madupu et al (2012). Nucleic Acids Research, 10.1093/nar/gkr1133.

The carbohydrate-active enzymes database (CAZy) in 2013.
V. Lombard et al (2014). Nucleic Acids Research, 10.1093/nar/gkt1178.

The Transporter Classification Database (TCDB): recent advances
M. H. Saier, Jr. et al (2016). Nucleic Acids Research, 10.1093/nar/gkv1103.

REBASE - a database for DNA restriction and modification: enzymes, genes and genomes.
R. J. Roberts et al (2015). Nucleic Acids Research, 10.1093/nar/gku1046.

Deep annotation of protein function across diverse bacteria from mutant phenotypes.
M. N. Price et al (2016). bioRxiv, 10.1101/072470.

by Morgan Price, Arkin group
Lawrence Berkeley National Laboratory